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The additive genetic effect for crossbred animals in formula (2) can then be expressed as c A B = c A B (A ) + c A B (B ), where c A B (A ) = 0.5 Z ~ A B, A c A + Φ A B (A ), c A B (B ) = 0.5 Z ~ A B, B c B + Φ A B (B ), and c A B (A ) and c A B (B ) are independent, i.e. the genetic effects for crossbred animals is split into two breed of origin genetic effects.
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