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On the lower side the midrib is often prominent.
In Actinostachys, or the ray ferns (16 species), the blades are reduced to a very narrow strip on either side of the midrib.
The sori vary from bean-shaped to linear and in most genera are positioned along both sides of the midrib of the leaflets or leaf divisions.
A marked characteristic of the leaf is an areolated portion bounded by two longitudinal curved lines, one on each side of the midrib, and more conspicuous on the under face of the leaf.
The glossy, bright green leaves, about 23 cm (9 inches) long and narrow toward the base, are divided almost to the midrib into seven or nine lobes; they turn wine red in autumn.
The black lines demarcate the angle of the midrib.
The leaves sometimes developed chlorosis along the midrib (Fig. 2b f).
Vascular tissue in the midrib is bicollateral type (Fig. 5a f).
The green paint was sampled from one of the leaves, close to the midrib (Fig. 1).
The result showed that the resistance-activating phenomenon was not able to move sideward across the midrib.
In the leaves, druse crystals were distributed either around the midrib or along the minor veins near xylem vessels.
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