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The functional morphology of the postcranial remains indicate that these elements come from animals that likely utilized terrestrial substrates, but they remain difficult to definitively attribute to Gorgopithecus, Theropithecus, or Papio given the absence of associated skeletons.
In the wild, Cebus monkeys utilize both arboreal and terrestrial substrates in their locomotor repertoire [39].
Amphibians deposit their eggs on a range of aquatic and terrestrial substrates [ 49, 67, 68], and previous studies have shown that sperm morphology can depend on spawning location [ 11, 69].
More firm conclusions in this regard will require studies of agonism in additional taxa, ideally including noncatarrhine primates that spend considerable time on the ground, as well as the comparison of rates of agonism on arboreal and terrestrial substrates in populations that spend a large proportion of time both in the trees and on the ground (Hill and Okayasu 1995).
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Furthermore, the continuum of benthic, near-shore, and fully terrestrial substrate interactions that many of these species engage in create a variety of impressions that, once preserved, would be equally difficult to match to environment.
Soil is the most ubiquitous terrestrial substrate, and in terrestrial ecosystems a rich biodiversity is found in soils and among surface litter, typically exceeding the biodiversity found above ground level [ 23, 24].
Because in many species agonism often occurs between coalitions, such coalition formation may be less likely if an individual has to move quickly through an arboreal environment than if it has only to move along a terrestrial substrate.
These findings suggest the presence of an enriched and varied array of carbohydrate uptake and metabolism mechanisms that can better facilitate the terrestrial substrate of L. rhinocerotis because such a lifestyle results in exposure to more diverse carbohydrate forms and sources as opposed to the wood saprophytes.
With the objective of in situ resource utilization to conserve energy and to limit transport costs, native materials mined on Moon or Mars are of primary interest for plant growth media in a future outpost, while terrestrial porous substrates with optimal growth media characteristics will be useful for onboard plant growth during space missions.
This paper summarizes recent advances in the understanding of the exchange of Hg between the atmosphere and natural terrestrial surfaces including substrates (soil, rocks, litter-covered surfaces and weathered lithological material) and foliage.
Pinnipeds also use a number of terrestrial habitats and substrates, both continental and island.
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