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The equilibrium was probably tipped by a highly successful breeding season for the southern couple.
These are the only known successful breeding attempts of the species in the US.
The park's successful breeding program with the white rhinoceros has led to a research program in conjunction with the San Diego Zoo.
The zoo also maintains successful breeding groups of gorillas, tigers, giraffes, hippopotamuses, bonobos (pygmy chimpanzees), Galapagos tortoises, and a wide variety of other animals.
First successful breeding happened in 1981 and from this time on regularly.
There is now a successful breeding population in the wild.
As pigs are polytocous species, litter size is a direct measure of efficient fertilization and successful breeding.
"These [baby Echidnas] represent the first successful breeding from zoo-born Echidnas and have shown that echidnas breed at a younger age than previously thought," writes Zooborns.com.
A succession of ospreys have subsequently taken up residence, culminating in successful breeding in 2011.
The fighting is believed to be necessary for successful breeding.
And last year was the most successful breeding year ever.
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