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In water exceeding the Dutch national pollution limit, just 17 species were found on average, whereas 52 species were found in cleaner water.
Working within a relatively small range, this and related species were found at several locations.
This makes it unlikely that all species were found and additional records should be expected.
This article was edited on 2 May: 52 species were found in cleaner water, not 50.
Individuals of the new species were found on all parts of the whale remains apart from those buried in sediment.
Twenty-eight other species were found, including thrushes, cuckoos and one green heron.
His achievements were many and varied, those around him recalling his enthusiasm and energy as new sites and species were found.
The new species were found by scientists, teachers and Indigenous rangers during a 10-day journey to the largely unsurveyed area.
During the second period up to 22 species were found.
Nine bacterial species were found in carp intestine.
A total of 223 introduced parasite species were found.
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