In contrast, sh4 explains less than 5% percentage of these variations when compared the degree of seed shattering between rice easy-shattering landraces and some accessions of O. rufipogon (Fig. 1).
Since sh4 is the result of the same mutation in indica and japonica rice, this mutation must have preceded mutation and divergence at the qSH1 locus that distinguishes the ease of shattering between these varietal groups.
The main QTL (qSH1) that distinguishes the degree of shattering between indica and japonica varieties explains 69% of the difference in these two groups of non-shattering rice [31].
However, recent analysis of the major gene that is responsible for the difference in degree of shattering between rice and wild rice has revealed that it is the same mutation in indica and japonica rice, which is not compatible with a dual origin of domesticated rice.
sh4 is the key shattering gene that distinguishes cultivated from wild rice, while the qSH1 gene controls the difference in the degree of shattering between some indica and japonica varieties of rice.
Others, including Harold Clurman and Robert Brustein, described the show -- also noted for its inclusion of live jazz music -- as a seminal work that shattered barriers between actors and audience.
Bloodshed had been limited compared with the fighting in other Libyan towns and cities - a total of four people had been killed - but the abuse of power by a minority of the town's Tuareg had shattered relations between the two populations.
But I am asking Mr Putin, would you remain?" The two leaders did not meet on Monday at the summit which Erdogan had previously described as an opportunity to mend shattered ties between the two countries.
By week's end, the earthquake had shattered any divide between those born in Haiti and those in Brooklyn.
For almost four decades, until the Council of Constance, unity was shattered by rivalries between popes and antipopes one French, one Italian, and later a third one, also Italian.
