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Two of these subjects (SL, LG) were additionally unable to perform search movements to detect single letters and isolated words initially projected within the scotoma, and to relocate the projected image onto a healthy peripheral area.
Assuming that specialist butterflies perceive their environment as fragmented (i.e. individuals are not able to perceive other suitable habitat patches when they leave their current patch), we hypothesize that dispersal movements in the matrix are relatively straight whereas search movements when foraging within habitat are rather tortuous.
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It may be that individuals in poor condition at eclosion invested more energy in search movement; this hypothesis could be tested by assessing metabolic rates or foraging behaviour under various conditions.
If a female loses an egg sac, she will make searching movements and may pick up a pebble or a piece of paper and attach it to the spinnerets.
Antennae of males of a domestic roach, Periplaneta americana, have specialized sense organs that detect the odour of female P. americana pheromones; upon detection of the odour, the male initiates searching movements, first with the antennae, then with the palpi.
The similar shape observed in all distributions suggests that identical mechanisms control the amplitude of the gaze shift, regardless of the systematic or random aspects of the searching movements on the 5-2 landices, and the distinctive features of the underlying arrangement of distractors that compose the "Where's Wally?" landscapes.
More recent versions of Walknet allow for searching movements when the agent is stepping into a hole.
Our results support the mounting evidence that dispersal in fragmented landscapes evolved towards the use of specific movement behaviour, different from explorative searching movements within habitat.
Indeed, our finding that dispersal movements in the matrix are straighter than resource searching movements within habitat supports the prediction of simulation studies that adopting straight movements for dispersal reduces its costs in fragmented landscapes.
More recent versions of Walknet allow for continuation of the swing movement by searching movements when the agent is stepping into a hole (not depicted in Fig. 5, but see Dürr 2001; Bläsing 2006).
In related studies Matheson and Dürr (2003) describe leg movements in locust where the hind leg performs searching movements targeted to different positions on the wing defined by a tactile stimulus.
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