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These signs can be summarized as cold temperatures and scarce grass in summer and heavy snowfall in winter (Figure 4).
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When grass is scarce or of too poor quality, buffaloes will browse woody vegetation.
Germany was deadly cold this time of year, the trees slick with ice, the grass so scarce it seemed the whole country had been poured in cement.
* If grass is scarce in June (TN21).
* If grass is scarce in summer, drought disaster may occur in winter (TN22).
Other work (Veblen 2008b) suggests that glades may serve as important wild herbivore foraging areas during the dry season, when grass is scarce in non-glade areas (sensu Ilius and O'Connor).
When grass is scarce, it will also eat the foliage of shrubs and trees.
Black wildebeest are predominantly grazers, preferring short grasses but also feeding on other herbs and shrubs, especially when grass is scarce.
They will accept forbs and dwarf shrubs, but only during dry periods when grass is scarce".
In comparison with the roan antelope, the hartebeest is better at procuring and chewing the scarce regrowth of perennial grasses at times when forage is least available.
The southward journey back to summer dry season camping areas starts mid-September to early October and has to be done rapidly, since water and grass become scarcer.
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