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Maestre, F. T., Castillo-Monroy, A. P., Bowker, M. A. & Ochoa-Hueso, R. Species richness effects on ecosystem multifunctionality depend on evenness, composition and spatial pattern.
Soil moisture appeared to be the major driver of plant and litter richness effects in our experiment.
Only species richness effects were tested in analyses of L. autumnalis because of low number of replicates.
Additionally, species richness effects can strengthen through time [42], an our experiments lasted only 7 10 months.
However, we saw significant herbivore richness effects on established benthic communities during these experiments and over this same time interval [16].
For time series data on percent cover, we used data from the final sampling period to test for herbivore richness effects.
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This was largely consistent in time, despite the tendency of varying plant species richness effect on herbivore density (Table S4).
The single substrate treatments xylose and galactose were associated with cases furthest from those showing transgressive overyielding, driving part of the substrate richness effect in the previous model.
We suggest that this absence of a species richness effect is partly explained by the relatively high richness values considered in our study (4 to 16 species) while past evidence for positive effects of species richness on ecosystem processes have often been due to the weak performances of monocultures or very species poor communities [2].
In fact, overlooking the allelic richness effect in our experiment, would also suggest enhanced resistance with increasing genotypic richness similar to reports from previous studies [ 50, 51, 57].
We were unable to detect any such consumer-richness effects on primary succession in this study, suggesting that individual species rather than consumer richness determined the trajectory of algal colonization on new substrates.
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