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The first class period of the module then starts with an activity.
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In a hierarchical scheme, losing a grid module produces location ambiguities that increase in size with the period of the missing module.
We then fix this resolution as being behaviorally determined and minimize the number of required neurons while allowing the periods of the modules, and, crucially, the number of modules, to vary to achieve the minimum.
This prediction could be tested behaviorally by comparing discrimination thresholds for location to the period of the smallest module.
Our theory predicts that the period of the largest grid module and the number of modules will be correlated with homing range.
As the CKD module was completed at different times across institutions, we were also unable to administer a follow-up test at a more remote period after completion of the module.
10.7554/eLife.08362.008 Appendix figure 1. Encoding range can exceed the period of the largest grid module at a cost in the number of neurons.
Likewise, the Introduction should be edited to read: "minimize the number of neurons required to achieve the behaviourally necessary spatial resolution across a spatial range equal in size to the period of the largest grid module" or similar.
Here, we propose that the grid system provides a hierarchical representation of space where fine grids provide precise location and coarse grids resolve ambiguity, and that the grids are organized to minimize the number of neurons required to achieve the behaviorally necessary spatial resolution across a spatial range equal in size to the period of the largest grid module.
For example, the Abstract should be edited to read: "We propose that the grid system implements a hierarchical code for spatial location that economizes the number of neurons required to encode location with a given resolution across a spatial range equal in size to the period of the largest grid module" or similar.
Three online formative MCQ tests were offered during the learning period of Musculoskeletal module on the 2nd week, 3rd week and 5th week for the students of group E. Each online test consisted of 20 MCQs designed to match the style and difficulty of summative examination.
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Since I tried Ludwig back in 2017, I have been constantly using it in both editing and translation. Ever since, I suggest it to my translators at ProSciEditing.

Justyna Jupowicz-Kozak
CEO of Professional Science Editing for Scientists @ prosciediting.com