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Most complex behaviors engage the body of the animal as a whole, and thus are expressed in a mutually exclusive manner.
Rather, as is the case with most complex behaviors and disorders, genetic contributions to the risk for alcohol dependence likely reflect the aggregate impact of a multitude of genetic variants and gene networks.
Male mating is one of the most complex behaviors in nematodes [ 63, 64], and many factors can influence successful mating: male sexual drive, ability of the male to locate the hermaphrodite, ability of the male to insert its spicules into the vulva, and amount of sperm transferred in each mating.
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Of course, the most complex behavior ever witnessed in chimpanzees is a far cry from what human societies can accomplish.
Close inspection of a subset of our simulations confirmed that the most complex behavior occurs for r = 3, with many trajectories reaching periodic orbits of the fourth type, that is, of length >3, that appear to have very small basins of attraction.
Delphinids display the greatest encephalization and the most complex behavior among cetaceans [ 9, 10].
The tool set used to deconstruct the circuits in hermaphrodites can be applied to study the most complex behavior exhibited by the nematode, male mating.
Consequently, dendrobatids exhibit some of the most complex spatial behaviors among amphibians.
Dendrobatidae (dart-poison frogs) exhibit some of the most complex spatial behaviors among amphibians, such as territoriality and tadpole transport from terrestrial clutches to widely distributed deposition sites.
Most complex insect behaviors, including flight, are modulated by various monoamines and neuropeptides and in Drosophila, flight can be modulated by octopamine, serotonin and dopamine as well as several neuropeptides (Taghert and Nitabach, 2012; Sadaf and Hasan, 2014; Van Breugel et al., 2014).
Arguably the most complex motor behavior displayed by this organism, male mating can be separated into at least six sub-steps: response, backing, turning, vulva location, spicule insertion, and sperm transfer [ 3].
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