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In short, Riedl recognized that morphological characters evolve at different rates.
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Of the three tested character systems, the evolutionary rate of "structural morphology" was in all analyses proportional to that of the mtDNA (irrespective of the method used to reconstruct the branch lengths), and thus it could be assumed that structural morphological characters evolved at rates proportional to time.
By calculating the proportion of observed changes to minimum changes, we derived convergence estimates that indicate that the female morphological characters are evolving at only approximately 2/3 the rate of molecular data (Fem: Obs./Min = 1.84; CI = 0.54).
How do your characters evolve this season?
Note although the derivation of Equations 3– 8 above is presented for nucleotide characters, these equations are also applicable to amino acid characters by substituting an amino acid substitution rate matrix for the nucleotide substitution rate matrix Q in Equations 1 and 2, and could also be applied to morphological characters that evolve in accord with the Mk matrix [ 61, 62].
Derived characters are used to infer evolutionary relationships, as derived characters evolved after primitive characters.
Character mapping also can inform us about the order in which different characters evolved.
This pattern is reminiscent of the non-parsimonious evolution of its morphological traits, including its inner ear and vertebrae, which indicate that this group is an early-branching lineage that diverged before those characters evolved.
The belief that every morphological or physiological character evolves by natural selection is still advocated by ultra-Darwinians (Eldredge 1995), who over-emphasize its power on organismal features.
The results also reveal several new examples where the traditional classifications have been misled by convergently evolved morphological characters.
Based on an analysis of how morphological characters used in defining Begonia sections evolve across a chloroplast DNA phylogeny (Thomas et al., 2011), it has become clear that Begonia sect.
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