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He is discovering more subtle traits through experience.
Now more than ever before, we want less mechanical and more subtle traits from our employees--traits that are harder to elicit using our old levers and measurement tools.
Since these aberrations affect many genes, their consequences are likely to obscure allelic differences at a single locus, especially when these are more subtle as for complex traits, and can thus lead to false positives.
This suggests that also for this beneficial trait specific alleles could be identified that would allow to improve this trait in a more subtle way.
It has been proposed that the alleles underlying complex traits have more subtle effects on disease risk and are hence more likely to include variants that affect the gene function modestly (H irschhorn and D aly 2005).
Many mutations affect a given phenotypic trait in much more subtle ways, and this translates the fact that the mapping of genotypic into phenotypic effects is much more complex, involving more than just a few genes with major effects.
The introduction of such traits may have broader but more subtle effects on the ecology of the production system.
The second type of cultural group selection is more subtle because the faster growth of some cultural groups is a result of their cultural traits being copied by neighboring groups more than other groups' traits (e.g., Henrich 2004).
It's more subtle".
Mimes are more subtle.
It was more subtle.
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Since I tried Ludwig back in 2017, I have been constantly using it in both editing and translation. Ever since, I suggest it to my translators at ProSciEditing.

Justyna Jupowicz-Kozak
CEO of Professional Science Editing for Scientists @ prosciediting.com