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Memory decay is rapid at first and slower later-a feature that accounts for Jost's memory law: that old memories gain on newer ones with lapse of time.

On the other hand, the time scale of the memory decay is

First, error-driven memory formation is context dependent; second, memory decay is context independent.

In motor tasks, memory decay is typically associated with reduced force output [ 10, 11].

Therefore, in the absence of any noticeable forgetting, we believe that memory decay is an unlikely mechanism driving subjects' performance.

In contrast, memory decay is assumed to be context independent, so that movement in any context leads to uniform decay across all contexts.

Moreover, since both MB structure and memory decay were apparently spared in HS flies, we argue that normal KC projection and connectivity are critical for memory storage and retrieval.

In the first model, memory decay was assumed to be context independent, consistent with previous studies (Equation 1).

In both cases, the context-dependent pattern of memory decay was still observed (see Supplemental Experimental Procedures and Figure S3).

This indicates that memory decay was greatest during probe trials at the initial exposure orientation, with progressively less decay occurring at more distant probe orientations.

In a previous report, we found that memory decay was greater in the rural area than in the urban area [ 19].