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This observation has been explained by the theoretical prediction that sex-ratio distortion enables a parasite to maintain its prevalence through vertical transmission alone [ 9, 10].
The maximum level of virulence possible if a vertically transmitted feminiser is to maintain its prevalence depends, crucially upon the sex ratio produced by uninfected hosts (g) as well as upon the parasite's rates of vertical transmission and feminisation [ 44, 52, 55].
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