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Outcomes were measured at the client (referrals), line staff (attitudes), and organizational (linkage) levels.
The same correction was done for those parameters influenced or correlated with network size (as linkage levels for plants and pollinator species) and similar results were obtained when correcting for number of species in the communities instead of network size.
Within each island, there were differences among localities in linkage levels for both plants (Ln) and insects (Lm) (Table 3); they were significantly higher at Cardona in Menorca and at Windmill in Tenerife.
That link density was unaffected by the alien was probably related to the lower pollinator linkage levels found in the invaded bi-modal networks; that is, rather than allowing the establishment of new interactions between insects and native plants in the invaded sites, aliens usurped links among them.
Our prediction on the impact of the alien on linkage levels is thus only partly confirmed; Opuntia does not modify the number of insect species that native plants interact with, but it can promote changes in the number of plants that insects visit.
This apparent contradiction (changes in Lm but not in Ln) could be attributed to the lower number of plants compared to insects in the pollination networks and to the higher heterogeneity (and thus greater standard deviation) in the linkage levels of the former.
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Comparing islands, we found higher linkage level for pollinators in Tenerife than in Menorca, which may be due to the oceanic nature of the former.
However, differences in the plant linkage level were not significant; i.e. overall, plants were visited by similar numbers of insect species in invaded and non-invaded sites.
Average linkage level for plants in the community increases if the alien plant promotes facilitation over competition for pollinators (i.e. if it mediates new interactions between pollinators and native plants); likewise, linkage level for insects decreases if the alien competes with natives for pollinators.
In the Canarian locality of Ravine, the significantly higher pollinator linkage level observed in 2005 compared to 2006 at the invaded site might also be attributed to the drier weather during the second year (2005: mean spring temperature (MST) = 18.9°C, total spring precipitation (TSP) = 104.9 mm; 2006: MST = 19.9°C, TSP = 41.0 mm).
Addition of links is assumed to be independent of the pollinator's present linkage level.
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