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If A1 conveys a weak advantage on the island (a < a*), it will almost immediately go extinct when it initially arises on background B2.
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When a novel allele of a gene initially arises in a population, it is associated with a single haplotype.
While it is tempting to view this as a conflict between Athens and Jerusalem, the problem initially arises within the Greco-Roman tradition itself [see Rist 1994, pg. 268].
The identification of the cells of origin from which cancer initially arises is of great importance, both for our understanding of the basic biology of tumours and for the translation of this knowledge to the prevention, treatment, and precise prognosis of the human disease (Visvader, 2011).
Although the Mets say that Santana's elbow troubles are normal and did not develop until after he learned he would not pitch in the World Baseball Classic, it is possible that they initially arose because he pushed himself too quickly to be ready for the Classic, even though the Mets did not want him to participate.
Interest in Y. lipolytica initially arose from its uncommon physiological characteristics.
When reproductive isolation is complete or nearly complete, it becomes more difficult to determine how barriers to gene flow initially arose - although there is no guarantee that those same barriers will complete it.
However, individuals with autism typically do not exhibit the same degree of connectivity or synchronization as control subjects, and it is from this evidence that a number of under-connectivity theories initially arose [ 50].
Leaves initially arise from cell divisions in the shoot apical meristem.
Problems initially arose with the remaining flotilla participants last week when Audacity of Hope, a US-owned ship, was barred from leaving Athens for a Greek island after suggestions that it was not seaworthy.
Nyong'o's interest in the subject matter initially arose out of a personal relationship she had with an albino in her neighborhood, but was further fueled by the tragedies of widely publicized Albino killings in the country of Tanzania.
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