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In all analyses we constrained the age of the root to be between 300 Myr and 600 Myr ago.

In all analyses we used a fixed study area, a fixed set of environmental predictors and a fixed set of presence observations.

In all analyses we considered p < 0.05 as statistically significant.

In all analyses, we report population means±SE except when noted (SD = standard deviation).

In all analyses, we adjusted for age using age sets consisting of three sequential Manjaco age groups.

In all analyses we therefore used a laboratory determination whenever one was available and otherwise adjusted the field tester measurement using this correction factor.

In all analyses we also included sex and its interactions but these results are not related to out predictions and did not interfere with the observed patterns and therefore will not be reported.

To ensure that only effects on trait variability were considered, in all analyses we discarded any trait/marker pairs with even marginally (nominal p<0.01) significant association between marker genotype and trait median (see Materials and Methods).

In all analyses we used robust standard errors, taking account of possible clustering within siblings (including twins or higher order multiple births), to calculate p-values and 95% confidence intervals.

In all analyses we used current prices.

In all analyses we controlled for baseline values and clustering.

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