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Tarnita and Nowak replied in Nature, and then again on their Web site, making it clear that they were not trying to discredit Hamilton or disregard the importance of relatedness.
He says he and his colleagues do not negate the importance of relatedness in the evolution of eusociality, as the critics say.
Overall, phylogenetic relatedness explained a relatively small proportion of the variation in leaf out timing across our 1126 tree species, but the importance of relatedness varied across the phylogenetic levels.
We also explored the importance of relatedness across four different phylogenetic scales (species, genus, family and order) to examine the level at which relatedness has the strongest influence on leaf out timing.
The importance of relatedness between validation and reference animals was also evaluated for the different chips and methods.
While the importance of relatedness for the evolution of public-goods cooperation has been demonstrated in several recent empirical studies of microbes [ 6- 8], the effect of variation in the cost of cooperation remains less well explored.
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The relative importance of phylogenetic relatedness increases towards lower taxonomic levels.
Increasingly, however, the relative importance of genetic relatedness for the evolution of reproductive skew is being questioned [ 1, 6- 8].
Our study is an additional example putting forward the importance of considering relatedness in a sample before studying it.
Our results underscore the importance of factors besides relatedness as the ultimate explanation for patterns of policing behaviour.
At the ultimate level, our results emphasize the importance of factors besides relatedness in predicting the level of conflict within a colony.
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