"implicated as the causes" is an acceptable and usable phrase in written English.
It can be used when discussing or pointing out reasons for something, as in the example: Recent reports have implicated climate change and air pollution as the causes of the decline in bird populations.
Although prions are merely self-perpetuating proteins, they have been implicated as the cause of various diseases, including bovine spongiform encephalopathy ("mad cow disease"), and are suspected of playing a role in a number of other disorders.
However, complexation to the membrane coordination sites is implicated as the cause of modifications to the permeation properties of the membranes.
Breakdown of blood-retinal barrier and the role of several inflammatory and vasoactive peptides have been implicated as the cause of macular edema in JIA [13].
Scientists have created an artificial prion, a type of protein whose misshapen version is implicated as the cause of several fatal and infectious conditions, including "mad cow disease" and human Creutzfeldt-Jakob disease.
Here pressure is implicated as the cause of the need for adaptation, acclimation, or adjustment: "Environmental pressures include things such as climate and habitat that would cause an animal to need to change something about itself in order to better survive".
Genetic factors may also be implicated as the cause of AVN.
Carbon monoxide poisoning has also been implicated as the cause of apparent haunted houses; symptoms such as delirium and hallucinations have led people suffering poisoning to think they have seen ghosts or to believe their house is haunted.
While no gene has been implicated as the cause of DPB, mutation in a specific gene much more likely to occur in Europeans causes CF. This mutation in the CF-causing gene is not a factor in DPB, but a unique polymorphism (variation) in this gene is known to occur in many Asians not necessarily affected by either disease.
In 2001, its mutations were implicated as the cause of the Blepharophimosis Ptosis Epicanthus-inversus Syndrome (BPES; MIM 110100) in humans [1].
Substrates for the ubiquitin ligase activity of a number of TRIM proteins have been identified [10]; for example, TRIM18/Mid2, implicated as the cause of X-linked Opitz syndrome, degrades protein phosphatase 2A [12A.
