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The maternal optimum remains the same as multiple mating does not change her relatedness to offspring, and the same is true for matrigenes in the worker.
The worker's relatedness to the focal male being helped is the same as her relatedness to other coeval males being harmed, r M = r coM, so: or if the focal male suffers a cost to benefit coevals Table 6 shows the thresholds.
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Furthermore, we found no effect of polyandry on female fitness, regardless of her level of relatedness to her mates.
Second, her blood-relatedness seems to imply a special relationship at least similar to that of the fiduciary relationship owed by a physician to her patient.
In the absence of children, who are an important element for a woman's social standing, this woman valued the relatedness to her husband.
For the question of placing an imprint that acts through the daughter, the appropriate r F is the queen's relatedness to her own particular daughter, 1/2 while the appropriate r coF is the weighted average of the queen's relatedness to her own daughters and to the daughters of other queens, using (4).
Each female tested was given the opportunity to choose a mate among either two of her brothers or two unrelated males (Relatedness treatment).
If one group member unilaterally chooses a goal for the group to address, this can hinder his or her groupmates' work processes (Yamaguchi, 2001) and relatedness (Capdeferro & Romero, 2012).
None of the measures of relatedness between the female and her social mate (r), and inbreeding level of the male (IR) affected the female likelihood to engage in extra-pair fertilizations, since the model with the lowest AIC value was the null model (Table 3, n = 70).
When, for instance, Martin regrets that he lost touch with his stepmother after her divorce from Kingsley, he laments "our vanished relatedness, cancelled by law but not by feeling".
In haplodiploid species, optimal sex ratio further depends on the level of inbreeding, because inbreeding increases the relatedness asymmetry between a mother and her sons and daughters.
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