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The approximate average invasion probability π ˜ ¯ is obtained according to Equation 4, with q ^ B as in Equation 2. Formally, these approximations are justified if ξ ≪ 1 (File S1, sect. 4). Figure 2 suggests that the assumption of weak evolutionary forces is more crucial than ξ small and that if it is fulfilled, the approximations are very good (compare Figure 2, A D).
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