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Lambda is a function of the traits heritability and pedigree structure (expressed as a kinship matrix).
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Does the function of the trait matter?
To make matters even more complex, analyses of homology may also need to consider the function of the trait.
Making even a general differentiation between them depends on specifying the organisms of interest, the aspect of the trait being compared, and even the function of the trait at various points in its history.
From a scientific perspective, being able to explain how a particular trait of an organism (be it morphological, developmental, behavioral, or some aspect of its life history) affects the fitness of an individual involves formulating hypotheses about the "function" of the trait.
The expected density of traits is an increasing function of trait flexibility and a decreasing function of the trait's deleterious potential (risk level).
However, although this method informs the function of the trait, it does not say much about the genetic mechanisms that shape this trait.
Integrating evo-devo with ecology requires model systems that allow for tests of gene expression and function from the one hand, and for tests of the function of the trait on the other hand.
Other traits such as systolic and diastolic blood pressure show a trait variance that increases with the magnitude of the measurement, in which case weights can be estimated as a function of the trait.
The basic understanding of the functions of the traits and how it impacts the interaction of organisms with their environment allows the identification of gene functions that affect not only the phenotype but also which genes matter the most in a given ecological interaction.
The Panglossian paradigm serves as a metaphor for some biologists' adaptationist approach to evolution, where the presence of traits is explained through the useful function of those traits.
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