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Doing so, once soil model was analyzed with single stone column of diameter 1 m at center of model with lower permeability (equal to surrounding soil permeability) without considering drainage boundary in stone columns environment with the same acceleration time history of VELACS project.
Evidence for such a pattern is provided by five markers which were outliers in both comparisons across the drainage boundary but showed FST values around or below the average in the comparison between the two species in sympatry in the Rhone.
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Next using the same acceleration time history of VELACS project, the model was analyzed with stone columns group with diameter equal to 1 m and center to center distance equal to 3 m and lower permeability (equal to surrounding soil permeability) without considering drainage boundaries in columns environment.
The Colorado River basin shares drainage boundaries with many other major watersheds of North America.
This pattern would only be observed if overland dispersal across drainage boundaries were a dispersal mechanism in the group.
Beyond the Northern Rockies clade however, all other clades except the Western Great Basin (which consists of a single locality) also show closely related haplotypes that span drainage boundaries.
This suggests that in certain areas of the species distribution, dispersal via overland flight across drainage boundaries may be more common than dispersal via overland flight (or larval drift) between distant localities (100 500 km distant) on the same river.
Estimates of gene flow suggested that migration rates between localities in adjacent drainage boundaries may be comparable to, or higher than migration rates between geographically proximate localities with a direct hydrologic connection.
The flow network of the gas in the borehole and from each gas source is shown in Fig. 2, where point a is the drilling wellhead, point b is the junction of the borehole and the protected layer, point c is the gas drainage area boundary of the protected layer, point d is the drilling downhole, and point e is the gas drainage area boundary of the fissure zone.
The perimeter (P) is the total length of the drainage basin boundary.
We expect headwater dispersal to be of particular importance in areas where drainage basin boundaries fall within (or near) the elevational distribution of the species, as observed for the boundary between the Green River and Hoback River localities.
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