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Such coloration features are features of the actual leaf lamina but coloration simply of the costa base is mostly irrelevant to species distinction.
Based on our study, Acrophorus belongs to Dryopteris, and represents a specialized group within Dryopteris with round indusia and cordate scales at costa base.
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However, the transverse costa at the base of sternum A2 found in most Yponomeutoidea and proposed to be a possible groundplan autapomorphy for the superfamily [ 62] was also found to be present in Roeslerstammiidae (Gracillarioidea): Roeslerstammia (Fig. 6), Vanicela, Amphithera and the female of Thereutis.
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