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Possible actions include establishing or expanding well-managed habitat preserves in all core populations, moving individuals to promote genetic exchange, and strategically restoring sage habitat to link the now-isolated populations.
Our maps depicting core populations and tillage risk enable partners to target conservation in landscapes where pipits will benefit most.
It is a common assumption that environmental conditions experienced by marginal or disjunct populations are more stressful or different than those experienced by core populations (Lesica and Allendorf 1995; Hardie and Hutchings 2010; Hart et al. 2010).
The species complex appears to have radiated from an origin in eastern Victoria (Jones, 2002), with well-defined core populations (Figure 1), and extensive intergrades between species at the boundaries.
The behavioural differences in mating that we observed from what is typical of core populations may reflect local adaptation to marginal conditions and could benefit the species in the face of ongoing environmental change.
In comparison to core populations, peripheral populations have low density and recruitment, and are subject to different selective pressures, such as environmental conditions, food type and availability, predation, disease, etc., which may result in behavioural modifications to mating.
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Between 1970 and 1997 Frankfurt, the German financial center, saw its core population drop, while the less densely populated suburban periphery, now extending to as much as 50 to 80 kilometers away, expanded dramatically.
The city's core population, excluding the 1,440 inmates, dropped in the 1990's to 2,644.
In 2014 it attracted about 3m visitors from all over the world to a city whose core population is 500,000.
"We think we can address the challenges of this core population," Ms. Eggleston said, "but we need flexibility.
The Wanapum band's core population lives in a village called Priest Rapids, at the base of the Priest Rapids Dam.
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