The phrase "copulation by" can be considered correct and usable in certain contexts in written English.
Specifically, it is typically used in scientific or technical writing to describe the act of sexual reproduction or mating in animals. Example: "The study observed the behavior of birds during copulation by utilizing remote cameras." In this sentence, "copulation by" is used to indicate the method or means by which the birds engage in sexual reproduction.
The female invites copulation by tilting forward and squatting, and the male approaches and mounts, sometimes while waving his wings.
Representative David Bates, the Republican who filed the repeal bill, argues that homosexuality is a lifestyle choice, and he even included a sentence that says: "Children can only be conceived naturally through copulation by heterosexual couples".
A similar situation occurs in an Australian orchid, Cryptostylis leptochila, which bears a sufficient resemblance to the female of the ichneumon wasp Lissopimpla semipunctata to induce copulation by the male wasp.
Mr. Bates said Monday that he was working on ways to broaden the bill's support in both chambers, like changing or removing a sentence that states, "Children can only be conceived naturally through copulation by heterosexual couples".
Hirst's first major show in New York came in 1996, at the Gagosian gallery (plans for a 1995 show had to be abandoned because, in Tomkins's words, "the New York City Department of Health took issue with the sanitary arrangements for the main art work, a glass case in which cows were supposed to simulate copulation by means of a hydraulic apparatus").
Benefits of stabilizing the copulation and intromission are indicated as the driving force for the evolution of extremely long aedeagi, while the short aedeagi might have the advantage of freedom of movements facilitating the initiation of copulation by males.
What has only recently been discovered is that males in many species make subtle alterations to their ejaculates on a copulation-by-copulation basis, depending on their assessment of immediate sperm competition risk.
The latter finding is consistent with audience effects from non-alpha individuals found in rhesus macaques (Overduin - de Vries et al. 2012) and long-tailed macaques (Overduin - de Vries et al. 2013), and with disruptions of copulations by non-alpha macaques that outrank one of the copulating partners (Niemeyer and Anderson 1983; Niemeyer and Chamove 1983; Manson 1996).
According to a 2010 study, males achieve almost three extra copulations by tricking females into staying.
Although females are receptive towards extra-pair copulations, male mate-guarding greatly reduces the rate of successful female involvement in attempted extra-pair copulations by other males.
Similarly, the proportion of attempted copulations by non-musth males did not differ between the three conditions (mean ±sd copulations attempted by non-musth males: A = 0.37±0.5; B = 0.33±0.5; C = 0.35±0.5; χ2 = 0.07, df = 2, p = 0.97. Note: mean values are shown here to better illustrate the spread of proportions).
