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The easy answer is that it was probably a combination of forest loss and market hunting.
Forest size and the combination of forest size and % bean cover in the matrix explained 37.5% and 42.1% of the variation in parasitism, respectively.
The combination of forest inventory and satellite-derived landscape composition and structure provides otherwise unavailable information on regional forest conditions and enables investigation of the cumulative effects of forest management over time.
By using a combination of forest sub-class, structure and disturbance variables within the MaxEnt framework, distribution maps of suitable langur and gibbon sleeping, ranging and feeding habitats were produced.
Analysis using conditional inference trees suggested that a combination of forest composition, topography, and dispersal predicted the distribution of MPB infestation on the landscape and that the importance of these variables changed as the outbreak developed.
Reasons behind the extirpation of taxa are examined in detail, and include a combination of forest clearance and human activities, isolation of populations, lack of temporal continuity of habitats, edaphic and competition factors affecting distribution of host trees (particularly pine), lack of forest fires and a decline in open forest systems.
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From the 164 species collected, 37 were indicators for various combinations of forest type, harvesting treatment and retention level.
In the summer of 2013, 18 field plots (45 m by 45 m) were established representing different combinations of forest types, density and leaf area index (LAI) (Table 1).
Finally, we apply multiobjective optimization to find optimal combinations of forest management regimes that maximize harvest revenues and carbon storage/sequestration.
Therefore it is necessary to encourage those combinations of forest structure and urban morphologies that benefit both processes in order to reduce the urban heat island.
A general finding is that targeted combinations of forest stand management strategies can lead to a higher degree of goal fulfilment at the landscape level than current forest management practice.
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