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Cladophora, however, can also be easily removed from coarse substrates during periods of hydrologic instability.
Diversity and population size of fish species such as Labeo altivelis and benthic macroinvertebrates associated with coarse substrates can be greatly reduced if the substrates are covered with sand and silt.
Alterations in stream morphology and hydrology, added to the effects of global climate change, are responsible for this phenomenon, that is particularly pernicious in mountainous lotic systems naturally characterized by coarse substrates.
In some older individuals, and those that burrow frequently in coarse substrates, the shell may be smooth.
The combination of a higher tolerance for sulphidic environments and a preference for coarse substrates, likely makes this genus more successful at vents.
It prefers gravelly and coarse sandy plains and large wadis and, although the species is typical of hard coarse substrates, it sometimes penetrates some dune areas [ 98].
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Substrate coarseness score was positively related to increasing heterogeneity in flow velocity, with increased presence of coarse substrate indicative of greater heterogeneity in velocity.
Observed responses to drought were consistent with the following two hypotheses: (i) Areas of coarse substrate can act as disturbance refugia for plants during drought by helping preserve pockets of water under large clasts and within absorbent materials such as brick.
However, during the cold winter months in the northern parts of its range, the long-toed salamander burrows below the frost-line in a coarse substrate to hibernate in clusters of 8 14 individuals.
The linear relationship between the change in area of coarse substrate littoral area and exposed beach were developed for each taxon.
During the period of rapid growth and reproduction (May through October), high water levels will expand the productive area while lower water levels will contract the available shallow-water area with coarse substrate.
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