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Millikan considers a related objection to do with omnipresent beneficial background conditions, the prima facie worry being whether her theory excludes them.
If a proportion of migrants carries a beneficial background allele, the mutation is less likely to become established.
On the other hand, when A1 initially occurs on the beneficial background, recombination is always deleterious, as it breaks up the fittest haplotype on the island (A1 B1).
In practice, however, their application requires accurate estimates of the recombination map, the selective advantage of the beneficial background allele, and the actual migration rate.
The most important, and perhaps surprising, effect is that the average invasion probability decreases with increasing continental frequency qc of the beneficial background allele B1.
Figure 2 shows this classification: a target mutation interacts with a deleterious background interfering mutation, with a beneficial background interfering mutation, and with a beneficial future interfering mutation.
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The SNS provides an intense flux of neutrinos in the few tens-of-MeV range, with a sharply-pulsed timing structure that is beneficial for background rejection.
The Spallation Neutron Source (SNS) at Oak Ridge National Laboratory, Tennessee, provides an intense flux of neutrinos in the few tens-of-MeV range, with a sharply-pulsed timing structure that is beneficial for background rejection.
Sign epistasis - which occurs when mutations are beneficial in one background and deleterious in other backgrounds - introduces particularly strong constraints.
The results also demonstrated that it is both necessary and beneficial to design background music for video games in such a way that it matches the action taking place in the game.
Gould says he has found his broad science background beneficial to his patent work with the pharmaceutical, food science, and biotechnology industries.
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