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"Because he succeeded in becoming an independent production where everybody has become answerable only to himself, the chain of command had been reduced to two people – Russell and Lesley," he added.
The question of how much genetic differentiation occurs during speciation has become answerable only with the relatively recent development of appropriate methods for comparing genes of different species.
An example of this is the academy programme, whereby schools become answerable only to central government and not to locally elected representatives.
Once you're able to index videos, pictures and live content in the same way as text, that "why" question that Rabkin hinted at starts to become answerable.
As genome sequences for Latrodectus and related theridiid species become available, these questions regarding the evolutionary origin of latrotoxins may become answerable.
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Departments also became answerable to citizens.
They were rewarded by a rise in rank and became answerable for their actions to the tsar alone.
At the same time, he warned the new system of 20,000 schools enjoying freedoms might unwittingly lead to centralisation as they became answerable only to Whitehall.
As Twitter prepares to IPO, and becomes answerable to a new influx of investors, it's inevitable that it's going to have to find more and more ways to make its service more mainstream.
Sustainability became answerable to big business.
Meanwhile, Rahul Gandhi, the son of Sonia Gandhi, who leads the governing Indian National Congress Party, broke his conspicuous silence on the issue by delivering a parliamentary address in which he recommended that any new anticorruption agency become a constitutional body answerable to Parliament.
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Justyna Jupowicz-Kozak
CEO of Professional Science Editing for Scientists @ prosciediting.com