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The tunnel system was expanded and the rock became a keystone in the defence of shipping routes to the Mediterranean.
The resulting book won the approval of both Braque and Picasso and became a keystone in Golding's life.
Sonic Youth grew out of a rowdy, close-knit world and became a keystone in New York's music scene but Moore felt the band was running out of road after 30 years and 15 albums.
When she played Mrs. Pynchon, the publisher of The Los Angeles Tribune on "Lou Grant," the role became a keystone of her career; for it she won four Emmy awards.
(Jon Pareles) DANNY AND DUSTY (Saturday) Two decades ago a couple of busy musicians, Dan Stuart and Steve Wynn, took 36 hours away from their regular bands to record a twangy, boozy album, "The Lost Weekend," which became a keystone of alt-country.
Inviting fans to fire-breathe, stilt-walk and generally burlesque it up at their early shows, they grew a wing called the Dirty Business Brigade to co-ordinate the performers and became a keystone of the burgeoning dark cabaret movement, a parade of theatrical freaks, mimes and Moriartys.
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But in the shifting priorities under the Obama administration, the station has become a keystone.
Such a mega-structure would become a keystone for regional co-operation.It will almost certainly never be built.
In a lyrical memory that becomes a keystone for the book, O'Neill recalls the evening her father taught her to cast a fishing line, pretending that the street was a trout-rich stream, goading her to swing the rod again and again as the streetlights came on, repeating: "Keep your eyes on the river, Molly.
A second issue is that a stone does not become a keystone until it is added to the rest of the assembled arch, and serving this role cannot be the reason it is maintained until that point.
See research article http://www.biomedcentral.com/1741-7007/10/59 Caelegansdithe elegans, the free-living nematode tamed as a new model organism by Sydney Brenner in the 1960s [ 1], has become a keystone species in the ecology of scientific knowledge.
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Justyna Jupowicz-Kozak
CEO of Professional Science Editing for Scientists @ prosciediting.com