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When the G tree is correct, then 1 and 2 are sister taxa and UF2 has expected value zero as will be demonstrated with reference to figure 3 A. We know from the two-taxon EP invariants, that a transversion must occur on either branch 1 or 2 in order to produce the Pu1Py2 pattern.
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The use of such indicators to reconstruct past instances and patterns of erosion is demonstrated with reference mainly to 50 British and Irish lake sites and especially those which have produced signals of erosion in the form of accelerated sediment accumulation and inversed or reversed sequences of radiocarbon dates during the mid and late Holocene.
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