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Previous research has shown that young Agathis trees can be damaged by full sunlight (Whitmore 1977), and along with other temperate rainforest species, Agathis have been shown to have lower chlorophyll concentrations in full sunlight compared with trees grown in medium and heavy shade (Langenheim et al. 1984; Read 1985).
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CEO of Professional Science Editing for Scientists @ prosciediting.com