In that paper, authors Paul Ehrlich and Peter Raven proposed that organisms that are constrained by other organisms, such as prey species, will develop novel defences to escape those constraints, and this "escape" then allows them to "radiate" into new species.
Additionally, luck-egalitarians may simply concede that the pursuit of the luck-egalitarian ideal is constrained by other ideals, including that of equal respect.
It may be that organizational form D is not available; another possibility is that even if all four organizational forms are available the selection of organizational form D is constrained by other factors.
The age of the Ongeluk Formation had long been under debate because two possible depositional ages (2.2 and 2.4 Ga) were constrained by other formations stratigraphically close to the Ongeluk Formation (Bau et al. 1999; Cornell et al. 1996; Dorland 2004).
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We can not completely exclude effects of heterosis as seed set in fully receptive pistils may have been constrained by other factors, e.g. nutrients.
It might be that the 12 per cent of children who do not attend basic schooling are constrained by other factors, including access to schools and cultural factors, which the Programme has not addressed.
A process evaluation 28 carried out in connection with a trial of educational visits to encourage general practitioners to follow prescribing guidelines 29 found that the visits were well received and recall of the guidelines was good, yet there was little change in prescribing behaviour, which was constrained by other factors such as patients' preferences and local hospital policy.
In particular, further work is needed to understand how phenology correlates with and is constrained by other traits, whether this varies between different climate regimes, functional groups and clades, and whether non-native species appear to face the same constraints to their phenologies as native species.
