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The phrase "are accumulated when the" is correct and usable in written English.
It can be used in contexts where you are discussing the process of gathering or collecting something under certain conditions or events.
Example: "The resources are accumulated when the project reaches its halfway point."
Alternatives: "are gathered when the" or "are collected when the".
Exact(1)
As SSCs that express neither pMad nor Bam are accumulated when the ecdysone pathway is perturbed it suggests that there should be an alternative mechanism of Bam regulation.
Similar(59)
Moreover, the losses had been accumulated when the Government - under pressure to maintain an aircraft manufacturing industry here - had pumped $650 million (Canadian) in investment grants into the development of the four-engine Dash-8.
The highest TFA content of 43.62 mg/g was accumulated when the aeration rate was set at 1.5 L/min.
These supported the result that starch was accumulated when the paper mulberry exposed to cold stress.
Aberrant gene expression may have been accumulated when they reach the haploid phase (i.e., in spermatids).
When ethanol was produced during growth on glucose or was accumulated when using ethanol or the mixed-substrate containing ethanol as a feed medium, the volumetric productivity did not improve.
On the other hand, more biomass was accumulated when ORP was controlled at −100 mV during ethanol fermentation with media containing 201 ± 3.1 and 252 ± 2.9 g/L glucose.
Otherwise, fermentations were categorized into 'abnormal.' Glutamate fermentation was non-growth associated, and glutamate was accumulated when cell growth had almost ceased (after 10 h).
HIF is accumulated when oxygen is limited [ 62, 63].
Up to 164 mM ethanol was accumulated when Thermoanaerobacter sp. DIB 004G was grown on glucose with pH control.
Ultra violet radiation leads to accumulation of phytoalexins (PA) in rice (Oryza sativa) which are typically accumulated when the plants are infected with rice blast pathogen Magnaporthe oryzae.
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Since I tried Ludwig back in 2017, I have been constantly using it in both editing and translation. Ever since, I suggest it to my translators at ProSciEditing.

Justyna Jupowicz-Kozak
CEO of Professional Science Editing for Scientists @ prosciediting.com