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The phrase "also characterized by substantial" is correct and usable in written English.
It can be used when describing additional features or qualities of a subject, particularly in formal or academic writing.
Example: "The study revealed that the species is also characterized by substantial adaptability to changing environments."
Alternatives: "also defined by significant" or "additionally marked by considerable".
Exact(1)
As a result, viral load was also characterized by substantial variation across pigs (Table 2).
Similar(59)
Most of the rest of the tree is also characterized by a substantial frequency of gene duplications and losses, and overall the family has few orthologs.
After isolation and expansion, cells were identified as adherent mesenchymal cells with substantial proliferation potential in vitro, and were also characterized by flow cytometry.
The late infancy period is also characterized by increasing infant exploratory behavior, fecal oral exposures, and diarrheal episodes, which will result in substantial shifts in the microbiota.
Still, 10 years ago California chardonnay was also characterized by ponderous extravagance.
And chlamydia, also characterized by burning with urination, is increasingly being screened in other ways.
The 18th century was also characterized by great social changes.
They are also characterized by a rare musicality and harmony.
The beads were also characterized by SEM and FTIR.
The macromolecules were also characterized by SEM, FTIR and DSC.
However, species are also characterized by persistent differences.
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Since I tried Ludwig back in 2017, I have been constantly using it in both editing and translation. Ever since, I suggest it to my translators at ProSciEditing.

Justyna Jupowicz-Kozak
CEO of Professional Science Editing for Scientists @ prosciediting.com