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Consequently, individual age was used to determine when individuals should be removed from the sexual network and was the basis for other age-specific traits.
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Selection against mortality declines with age, and it follows that age-specific trait values will decline with age as well.
We then standardised our age-specific trait means and tested whether age trajectories could be meaningfully grouped according to sex or the type of trait.
The response variable in these comprised all age-specific predictions from all traits, with explanatory variables coding either the age or the group of that trait in each model.
Relatively little is known about age-specific changes in functional traits and how demographic and functional senescence are genetically (co regulated.
Means and standard errors (in parentheses) were obtained by averaging over the age-specific estimates for the trait type.
We initially sought to calculate age-specific predictions for each trait in both males and females (where appropriate), whilst accounting for individual and annual variation and selective disappearance effects.
We applied generalised additive models (GAMs) to the age-specific predictions of the remaining traits from our GLMMs to test whether groups of traits identified a priori, showed similar or different senescence patterns (following Nussey et al., 2009).
Study of inbreeding effects in specific traits, such as age-specific mortality and life span, provide a good starting point, as a limited set of genes is expected to be involved.
Although age-specific genetic covariance matrices among traits have been estimated in laboratory model systems (e.g. Tatar et al., 1996), we are not aware of any such estimates from the wild (Charmantier et al., 2014).
The age-specific predicted means of all traits were then standardised to be on comparable scales by subtracting the age-specific predicted mean at age 4 for each sex from the age-specific predicted mean at each age and then dividing this by the range of age-specific predicted means from age 4 to 13 in females and 4 to 9 in males.
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