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The cytoskeleton of non-transfected BE cells was organized in the form of a cortical ring under the cellular membrane and actin meshwork in the cell body, with clearly defined lamellipodia, invadopodia and many subtle protrusions.
A subtle signal?
Iranians are a subtle people.
A subtle or obvious message?
It is a subtle war.
This is a subtle concept.
A subtle underlying levity prevailed.
A subtle portent lurks behind.
It's a subtle thing.
Again, a subtle lesson.
Wisconsin cheese adds a subtle locavore touch.
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Since I tried Ludwig back in 2017, I have been constantly using it in both editing and translation. Ever since, I suggest it to my translators at ProSciEditing.

Justyna Jupowicz-Kozak
CEO of Professional Science Editing for Scientists @ prosciediting.com