The British operational commander, Brig.
Gen. Benjamin C. Freakley, the operational commander of the United States-led coalition force, said in the statement.
He continued to be the terror network's operational commander, using couriers, e-mail, and coded telephone messages to maintain communications.
Zarqawi is more a ruthless operational commander, putting strategy into action: a thug rather than a thinker.
Mr. Zadran served as a top aide to Sirajuddin Haqqani, the network's operational commander and the son of the group's patriarch, Jalaluddin Haqqani.
Over the past two and a half years, he said, Al Qaeda's leadership has been "decimated," and virtually every affiliate has lost a top leader or operational commander.
Abdelhamid Abaaoud, 28 The presumed operational commander of the attacks died in a hail of 5,000 police bullets in Saint-Denis on Thursday.
The arrest of the Taliban's top operational commander, Abdul Ghani Baradar, in January has not led to a broader crackdown against the Afghan insurgents.
In a news release, the NATO mission's operational commander, Lt. Gen. Charles Bouchard, said he was aware of the reports of Qaddafi family deaths but called them unconfirmed.
Zaki ur-Rehman Lakhvi, the operational commander accused of masterminding the assault, is in jail, but efforts to prosecute him have been characterized by an opaque legal process.
He later joined the National Liberation army in the Constantine region of north-eastern Algeria, and served as operational commander in various provinces.
