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He is the father of nearly all the bears born in the mountains since 1997.
Precisely 17 more are known from zoos, the result of bears born to polar bears and brown bears kept together in enclosures, which subsequently mated.
Sonne's team gathered samples from the penile bones of 279 polar bears born between 1990 and 2000.
Bears born or raised in captivity have lost their fear of people or never develop fear of humans unless special measures are taken to reduce human contact or access to caregivers.
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When I was growing up, millions in Britain swooned over Brumas - a polar bear born in Regent's Park Zoo, London, in 1949 - the first bear to be reared in captivity in the UK.
Even a fifth generation polar bear born in a zoo is a wild animal".
My father, whose identical name I bear, was born in the small fishing village of Polruan in the Fowey Harbour.
"Regret you did not inform us bear wanted for work in extreme North, bear born near Hamburg, not accustomed to cold weather".
That includes a Disney tree (with all the classic characters dressed in Santa garb) and a polar bear tree he created four years ago to mark the birth of Knut, a bear born in captivity at the Berlin Zoo.
Are some bears just born to be bad?
Some skeptics point out that polar bears are born swimmers and that the worries of environmentalists are therefore overdone.
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Since I tried Ludwig back in 2017, I have been constantly using it in both editing and translation. Ever since, I suggest it to my translators at ProSciEditing.

Justyna Jupowicz-Kozak
CEO of Professional Science Editing for Scientists @ prosciediting.com